Billy Vunipola beat Alex Dombrandt to the sole specialist number eight spot in England's Rugby World Cup squad as Henry Slade also missed out.
Vunipola, 30, has not played for England so far this year and has carried an injury into this year's training camp, with brother Mako out of the squad because he failed to recover from a back problem.
But Vunipola has seemingly recovered, and gets into the squad ahead of Dombrandt who occupied the eight jersey during this year's Six Nations Championship.
Slade's omission, behind soon-to-be Stade Francais player Joe Marchant, is the other major talking point from the squad, whilst youngster Henry Arundell gets the call ahead of more experienced wingers Jonny May and Joe Cokanasiga.
Jamie Blamire started Saturday's defeat to Wales, but he is out of the squad in favour of Saracens' Theo Dan – who was on the bench for that clash – and Marcus Smith at ten makes the squad as one of six half backs, along with fellow Harlequin Danny Care at scrum-half.
“Rugby World Cups involve unique challenges in terms of the demands on players, the need for squad flexibility and the limited number of squad places available,” said Borthwick.
“France 2023 is no different, but I am confident we have selected a squad of 33 that is well able to meet those challenges and leave us best placed to be at our most competitive in this exciting tournament.
“There have been a number of very difficult selection decisions to make, given the quality of our wider training squad. Some very good players have missed out on selection, such has been the competition for places over the last eight weeks and more of preparation.
“As Saturday's game in Cardiff proved, we have a series of demanding Test matches through August as we continue that preparation towards our opening Rugby World Cup pool game against an in-form Argentina side.”
Forwards: Ollie Chessum (Leicester), Dan Cole (Leicester), Tom Curry (Sale), Theo Dan (Saracens), Ben Earl (Saracens), Ellis Genge (Bristol), Jamie George (Saracens), Maro Itoje (Saracens), Courtney Lawes (Northampton), Lewis Ludlam (Northampton), Joe Marler (Harlequins), George Martin (Leicester), David Ribbans (Toulon), Bevan Rodd (Sale), Kyle Sinckler (Bristol), Will Stuart (Bath), Billy Vunipola (Saracens), Jack Walker (Harlequins), Jack Willis (Toulouse).
Backs: Henry Arundell (Racing 92), Danny Care (Harlequins), Elliot Daly (Saracens), Owen Farrell (Saracens), George Ford (Sale), Ollie Lawrence (Bath), Max Malins (Bristol), Joe Marchant (Stade Francais), Marcus Smith (Harlequins), Freddie Steward (Leicester), Manu Tuilagi (Sale), Jack van Poortvliet (Leicester), Anthony Watson (unattached), Ben Youngs (Leicester).
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